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2023
Journal article
Hwiccewyrm trispiculum gen. et sp. nov., a new leptopleuronine procolophonid from the Late Triassic of southwest England
The fissure fill localities of southwest England and South Wales are well‐known for preserving rich assemblages of predominantly small‐bodied Late Triassic to Early Jurassic tetrapods, but many aspects of these assemblages remain contentious. The age of the Late Triassic fissures is disputed, with some lines of argument suggesting a latest...Butler, Richard J ; Meade, Luke E ; Cleary, Terri J ; McWhirter, Kai T ; Brown, Emily E …
Procolophonidae , phylogeny , Parareptilia , fissure fills , and Triassic
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2023
Journal article
New specimen and redescription of Anisodontosaurus greeri (Moenkopi Formation: Middle Triassic) and the spatiotemporal origins of Trilophosauridae
Anisodontosaurus greeri is an enigmatic small-bodied tetrapod with a heterodont dentition from the Holbrook Member of the Moenkopi Formation (Middle Triassic) of Arizona (U.S.A.). The evolutionary relationships of this taxon have long been debated and remain uncertain. Using micro-computed tomography (µCT) scans we redescribe the holotype of Anisodontosaurus greeri (UCMP...Foffa, Davide ; Nesbitt, Sterling J ; Kligman, Ben T ; Butler, Richard J ; Stocker, Michelle R
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2022
Journal article
Scleromochlus and the early evolution of Pterosauromorpha
Pterosaurs, the first vertebrates to evolve powered flight, were key components of Mesozoic terrestrial ecosystems from their sudden appearance in the Late Triassic until their demise at the end of the Cretaceous1,2,3,4,5,6. However, the origin and early evolution of pterosaurs are poorly understood owing to a substantial stratigraphic and morphological...Foffa, Davide ; Dunne, Emma M ; Nesbitt, Sterling J ; Butler, Richard J ; Fraser, Nicholas C …
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2021
Journal article
New species of mammaliaform and the cranium of Borealestes (Mammaliformes: Docodonta) from the Middle Jurassic of the British Isles
Docodonta are one of the earliest diverging groups of mammaliaforms, and their morphology provides key information on the transition between non-mammalian cynodonts and Mammalia. We describe the partial skulls of two docodontans Borealestes serendipitus and Borealestes cuillinensis sp. nov. from the Kilmaluag Formation (Middle Jurassic: Bathonian), Isle of Skye, Scotland....Panciroli, Elsa ; Benson, Roger B J ; Fernandez, Vincent ; Butler, Richard J ; Fraser, Nicholas C …
mammals, Scotland, Jurassic, Bathonian, Mesozoic, and Isle of Skye
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2020
Journal article
Revision of Erpetosuchus (Archosauria: Pseudosuchia) and new erpetosuchid material from the Late Triassic ‘Elgin Reptile’ fauna based on μCT scanning techniques
The Late Triassic fauna of the Lossiemouth Sandstone Formation (LSF) from the Elgin area, Scotland, has been pivotal in expanding our understanding of Triassic terrestrial tetrapods. Frustratingly, due to their odd preservation, interpretations of the Elgin Triassic specimens have relied on destructive moulding techniques, which only provide incomplete, and potentially...Foffa , Davide ; Butler, Richard J ; Nesbitt, Sterling J ; Walsh, Stig ; Barrett, Paul M …
systematics, Erpetosuchidae , Erpetosuchus granti , Leptopleuron , and anatomy
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2020
Journal article
Diverse vertebrate assemblage of the Kilmaluag Formation (Bathonian, Middle Jurassic) of Skye, Scotland
The Kilmaluag Formation on the Isle of Skye, Scotland, provides one of the richest Mesozoic vertebrate fossil assemblages in the UK, and is among the richest globally for Middle Jurassic tetrapods. Since its discovery in 1971, this assemblage has predominantly yielded small-bodied tetrapods, including salamanders, choristoderes, lepidosaurs, turtles, crocodylomorphs, pterosaurs,...Panciroli, Elsa ; Benson, Roger B J ; Walsh, Stig ; Butler, Richard J ; Castro, Tiago Andrade …
palaeontology , mammaliaforms , tetrapods, salamanders , Great Estuarine Group , and squamates
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2018
Journal article
New partial dentaries of amphitheriid mammal Palaeoxonodon ooliticus from Scotland, and posterior dentary morphology in early cladotherians
We describe two partial dentaries of mammals from the Middle Jurassic of Scotland. They belong to the early cladotherian Palaeoxonodon ooliticus. These dentaries comprise the first specimen of P. ooliticus ever found—although its significance was initially unrecognised so it remained undescribed until now—and the most recently discovered specimen, found during...Panciroli, Elsa ; Benson, Roger B J ; Butler, Richard J
UK, Mammalia, Scotland., Jurassic, Bathonian, Cladotheria, Kilmaluag Formation, and Palaeoxonodon ooliticus