True flies (Insecta: Diptera) from the late Eocene insect limestone (Bembridge Marls) of the Isle of Wight, England, UK

Abstract

• The Diptera fauna from the late Eocene of the Isle of Wight (Bembridge Marls) is studied including redescriptions of formerly described material. The fauna includes the following taxa: Anisopodidae – one species; Bibionidae – 11 species; Ceratopogonidae – one described and two unidentified species; Chironomidae – undetermined species of three subfamilies; Culicidae – four species; Cylindrotomidae – one species; Dixidae – one species; Keroplatidae – one described and four unidentified species; Limoniidae – 31 species; Mycetophilidae – 14 species; Psychodidae – two species; Scatopsidae – two species; Sciaridae – three species; Simuliidae – an unnamed species; Tipulidae – nine species. Diptera Brachycera: Agromyzidae – unidentified species; Anthomyzidae – two species; Asilidae – two species; Bombyliidae – one species; Chloropidae – one species; Dolichopodidae – unidentified species; Empididae – unidentified species; Ephydridae – two species; Heleomyzidae – unidentified species; Lauxaniidae – unidentified species; Otitidae – one species; Phoridae – unidentified species; Rhagionidae – two species; Stratiomyidae – two species; Syrphidae – unidentified species; Tabanidae – one species. Following new taxa are described: Leptotarsus (Longurio) fragmentatus KrzemiÅ„ski, n. sp., Leptotarsus (Longurio) wegiereki KrzemiÅ„ski, n. sp., Tipula (s. lato) anglicana KrzemiÅ„ski, n. sp. (Tipulidae); Cyttaromyia rossi KrzemiÅ„ski, n. sp. (Cylindrotomidae); Gurnardia corami KrzemiÅ„ski, n. gen, n. sp., Pilaria hooleyi KrzemiÅ„ski, n. sp., Pilaria volodii KrzemiÅ„ski, n. sp., Cheilotrichia (Cheilotrichia) duplicata KrzemiÅ„ski, n. sp., Cheilotrichia (Empeda) szwedoi KrzemiÅ„ski, n. sp., Symplecta (Psiloconopa) gurnetensis KrzemiÅ„ski, n. sp., Orimarga (Orimarga) lenae KrzemiÅ„ski, n. sp., Dicranomyia (Dicranomyia) azari KrzemiÅ„ski, n. sp., Dicranomyia (Dicranomyia) fasciata KrzemiÅ„ski, n. sp., Helius (Helius) edmundi KrzemiÅ„ski, n. sp., Helius (Helius) popovi KrzemiÅ„ski, n. sp., Dicranoptycha staryi KrzemiÅ„ski, n. sp. (Limoniidae); Wightipsychoda Azar, n. gen. (Psychodidae); Dilophus andrewrossi Nel, Colomb & Waller, n. sp., Bibiodes massiliensis Nel, Colomb & Waller, n. sp. (Bibionidae); Azana cockerelli Blagoderov, n. sp., Aglaomyia vectis Blagoderov, n. sp., Dziedzickia oligocenica Blagoderov, n. sp., Palaeoempalia saxea Blagoderov, n. sp., Mycomya hoolei Blagoderov, n. sp., Leia gurnardensis Blagoderov, n. sp. (Mycetophilidae); Regmoclemina haennii KrzemiÅ„ski, n. sp. (Scatopsidae); Sylvicola problematica KrzemiÅ„ski, n. sp. (Anisopodidae); Palaeoberidops barkeri Nicholson, n. gen., n. sp. (Stratiomyidae). The Diptera fauna and palaeohabitat of the Insect Bed is discussed.